1485–Present
Of or relating to the night; done, held, or occurring at night
1599–Present
Of an animal: active chiefly or exclusively at night
1840–Present
Of an eye: adapted for vision at night
1679–Present
Of a person: that engages in an activity or occupation by night; preferring to be active at night
Latin
Nocturnalis
Nocturnale
The frequency with which the variant noctural occurs suggests that it is not a straightforward transmission error; it may show remodelling after natural adj.